Titin in insect spermatocyte spindle fibers associates with microtubules, actin, myosin and the matrix proteins skeletor, megator and chromator.
Fabian L, Xia X, Venkitaramani DV, Johansen KM, Johansen J, Andrew DJ, Forer A. J Cell Sci. 2007 Jul 1; 120(Pt 13):2190-204.