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SH3D19 SH3 domain containing 19 [ Homo sapiens (human) ]

Gene ID: 152503, updated on 27-Nov-2024

Summary

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Official Symbol
SH3D19provided by HGNC
Official Full Name
SH3 domain containing 19provided by HGNC
Primary source
HGNC:HGNC:30418
See related
Ensembl:ENSG00000109686 MIM:608674; AllianceGenome:HGNC:30418
Gene type
protein coding
RefSeq status
REVIEWED
Organism
Homo sapiens
Lineage
Eukaryota; Metazoa; Chordata; Craniata; Vertebrata; Euteleostomi; Mammalia; Eutheria; Euarchontoglires; Primates; Haplorrhini; Catarrhini; Hominidae; Homo
Also known as
EBP; EVE1; Kryn; Eve-1; SH3P19
Summary
This gene encodes a multiple SH3 domain-containing protein, which interacts with other proteins, such as EBP and members of ADAM family, via the SH3 domains. This protein may be involved in suppression of Ras-induced cellular transformation and Ras-mediated activation of ELK1 by EBP, and regulation of ADAM proteins in the signaling of EGFR-ligand shedding. Alternatively spliced transcript variants encoding different isoforms have been found for this gene. [provided by RefSeq, Aug 2011]
Expression
Ubiquitous expression in fat (RPKM 43.0), gall bladder (RPKM 40.6) and 23 other tissues See more
Orthologs
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Genomic context

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See SH3D19 in Genome Data Viewer
Location:
4q31.3
Exon count:
24
Annotation release Status Assembly Chr Location
RS_2024_08 current GRCh38.p14 (GCF_000001405.40) 4 NC_000004.12 (151120281..151325605, complement)
RS_2024_08 current T2T-CHM13v2.0 (GCF_009914755.1) 4 NC_060928.1 (154444161..154649516, complement)
RS_2024_09 previous assembly GRCh37.p13 (GCF_000001405.25) 4 NC_000004.11 (152041433..152246757, complement)

Chromosome 4 - NC_000004.12Genomic Context describing neighboring genes Neighboring gene uncharacterized LOC107986196 Neighboring gene Sharpr-MPRA regulatory region 3738 Neighboring gene small nucleolar RNA, C/D box U73B Neighboring gene small nucleolar RNA, C/D box 73A Neighboring gene ribosomal protein S3A Neighboring gene uncharacterized LOC105377485 Neighboring gene uncharacterized LOC124900799 Neighboring gene CDK7 strongly-dependent group 2 enhancer GRCh37_chr4:152236121-152237320 Neighboring gene RNA, U6 small nuclear 1282, pseudogene Neighboring gene serine protease 48 Neighboring gene Sharpr-MPRA regulatory region 2958 Neighboring gene OCT4-NANOG-H3K27ac-H3K4me1 hESC enhancer GRCh37_chr4:152312507-152313010 Neighboring gene H3K27ac-H3K4me1 hESC enhancers GRCh37_chr4:152328899-152329598 and GRCh37_chr4:152329599-152330298 Neighboring gene FHIP1A divergent transcript Neighboring gene FHF complex subunit HOOK interacting protein 1A Neighboring gene OCT4-NANOG-H3K27ac hESC enhancer GRCh37_chr4:152405062-152405718 Neighboring gene OCT4-NANOG-H3K27ac hESC enhancer GRCh37_chr4:152405719-152406374 Neighboring gene H3K27ac hESC enhancer GRCh37_chr4:152406474-152407173 Neighboring gene H3K27ac hESC enhancer GRCh37_chr4:152407174-152407872 Neighboring gene NANOG-H3K27ac-H3K4me1 hESC enhancer GRCh37_chr4:152409975-152410608 Neighboring gene NANOG hESC enhancer GRCh37_chr4:152445639-152446203 Neighboring gene RN7SK pseudogene 35

Genomic regions, transcripts, and products

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Go to nucleotide: Graphics FASTA GenBank

Expression

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  • Project title: HPA RNA-seq normal tissues
  • Description: RNA-seq was performed of tissue samples from 95 human individuals representing 27 different tissues in order to determine tissue-specificity of all protein-coding genes
  • BioProject: PRJEB4337
  • Publication: PMID 24309898
  • Analysis date: Wed Apr 4 07:08:55 2018

Bibliography

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Related articles in PubMed

  1. Identification of novel Runx1 (AML1) translocation partner genes SH3D19, YTHDf2, and ZNF687 in acute myeloid leukemia. Nguyen TT, et al. Genes Chromosomes Cancer, 2006 Oct. PMID 16858696
  2. Gene expression of Sh3d19, a novel adaptor protein with five Src homology 3 domains, in anagen mouse hair follicles. Shimomura Y, et al. J Dermatol Sci, 2003 Feb. PMID 12615363
  3. Identification and characterization of EBP, a novel EEN binding protein that inhibits Ras signaling and is recruited into the nucleus by the MLL-EEN fusion protein. Yam JW, et al. Blood, 2004 Feb 15. PMID 14551139
  4. Comparative proteomics of clathrin-coated vesicles. Borner GH, et al. J Cell Biol, 2006 Nov 20. PMID 17116749, Free PMC Article
  5. ADAM binding protein Eve-1 is required for ectodomain shedding of epidermal growth factor receptor ligands. Tanaka M, et al. J Biol Chem, 2004 Oct 1. PMID 15280379

See all (46) citations in PubMed

GeneRIFs: Gene References Into Functions

What's a GeneRIF?
  1. occurrence of an unusual TG 3' splice site in intron 6
    Title: Violating the splicing rules: TG dinucleotides function as alternative 3' splice sites in U2-dependent introns.
  2. yeast two-hybrid screening using the cytoplasmic domain of ADAM12 as bait identified a protein designated Eve-1. Eve-1 plays a role in positively regulating the activity of ADAMs in the signaling of EGFR-ligand shedding.
    Title: ADAM binding protein Eve-1 is required for ectodomain shedding of epidermal growth factor receptor ligands.
  3. identified and characterized EBP, a novel EEN binding protein that interacts with the SH3 domain of EEN; normally expressed in cytoplasm but recruited to the nucleus by MLL-EEN [EBP]
    Title: Identification and characterization of EBP, a novel EEN binding protein that inhibits Ras signaling and is recruited into the nucleus by the MLL-EEN fusion protein.
Submit: New GeneRIF Correction

Phenotypes

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Review eQTL and phenotype association data in this region using PheGenI

Variation

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See variants in ClinVar

See studies and variants in dbVar

See Variation Viewer (GRCh37.p13)

See Variation Viewer (GRCh38)

Pathways from PubChem

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Interactions

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Products Interactant Other Gene Complex Source Pubs Description

General gene information

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Markers

Homology

Clone Names

  • MGC105136, MGC118910, MGC118911, MGC118912, MGC118913

Gene Ontology Provided by GOA

Function Evidence Code Pubs
enables proline-rich region binding IPI
Inferred from Physical Interaction
more info
 PubMed 
enables protein binding IPI
Inferred from Physical Interaction
more info
 PubMed 
Process Evidence Code Pubs
involved_in cytoskeleton organization IMP
Inferred from Mutant Phenotype
more info
 PubMed 
involved_in positive regulation of membrane protein ectodomain proteolysis IMP
Inferred from Mutant Phenotype
more info
 PubMed 
involved_in regulation of cell morphogenesis IMP
Inferred from Mutant Phenotype
more info
 PubMed 
Component Evidence Code Pubs
located_in cytosol IDA
Inferred from Direct Assay
more info
  
located_in cytosol TAS
Traceable Author Statement
more info
  
located_in nucleoplasm IDA
Inferred from Direct Assay
more info
  
located_in plasma membrane IDA
Inferred from Direct Assay
more info
 PubMed 

General protein information

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Preferred Names
SH3 domain-containing protein 19
Names
ADAM-binding protein Eve-1
EEN-binding protein
SH3 domain protein D19

NCBI Reference Sequences (RefSeq)

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RefSeqs maintained independently of Annotated Genomes

These reference sequences exist independently of genome builds. Explain

These reference sequences are curated independently of the genome annotation cycle, so their versions may not match the RefSeq versions in the current genome build. Identify version mismatches by comparing the version of the RefSeq in this section to the one reported in Genomic regions, transcripts, and products above.

Genomic

  1. NG_029721.2 RefSeqGene

    Range
    5000..210324
    Download
    GenBank, FASTA, Sequence Viewer (Graphics)

mRNA and Protein(s)

  1. NM_001009555.4 → NP_001009555.3  SH3 domain-containing protein 19 isoform 1

    See identical proteins and their annotated locations for NP_001009555.3

    Status: REVIEWED

    Description
    Transcript Variant: This variant (1) encodes the longest isoform (1, also known as Eve-1a).
    Source sequence(s)
    AC095055, AW263019, BC032404, BM670337, BM831166, BX647422, CD675494, DB277036, DB278138
    Consensus CDS
    CCDS34077.2
    UniProtKB/Swiss-Prot
    B7Z296, Q08EK1, Q32N10, Q5HYK7, Q5U3B8, Q86XB3, Q8N5E7, Q9UFC8
    Conserved Domains (6) summary
    cd11814
    Location:419 → 468
    SH3_Eve1_1; First Src homology 3 domain of ADAM-binding protein Eve-1
    cd11815
    Location:499 → 550
    SH3_Eve1_2; Second Src homology 3 domain of ADAM-binding protein Eve-1
    cd11816
    Location:575 → 625
    SH3_Eve1_3; Third Src homology 3 domain of ADAM-binding protein Eve-1
    cd11817
    Location:665 → 714
    SH3_Eve1_4; Fourth Src homology 3 domain of ADAM-binding protein Eve-1
    cd11818
    Location:734 → 783
    SH3_Eve1_5; Fifth Src homology 3 domain of ADAM-binding protein Eve-1
    PHA03247
    Location:29 → 369
    PHA03247; large tegument protein UL36; Provisional

  2. NM_001128923.2 → NP_001122395.1  SH3 domain-containing protein 19 isoform 2

    See identical proteins and their annotated locations for NP_001122395.1

    Status: REVIEWED

    Description
    Transcript Variant: This variant (2) lacks an in-frame coding exon compared to variant 1. This results in a shorter isoform (2, also known as Eve-1b) missing an internal protein segment compared to isoform 1. Variants 2 and 4 encode the same isoform.
    Source sequence(s)
    AC095055, AW263019, BC032404, BM670337, BM831166, CD675494, DB277036
    Consensus CDS
    CCDS47144.1
    UniProtKB/Swiss-Prot
    Q5HYK7
    Related
    ENSP00000387030.4, ENST00000409598.8
    Conserved Domains (6) summary
    cd11815
    Location:476 → 527
    SH3_Eve1_2; Second Src homology 3 domain of ADAM-binding protein Eve-1
    cd11816
    Location:552 → 602
    SH3_Eve1_3; Third Src homology 3 domain of ADAM-binding protein Eve-1
    cd11817
    Location:642 → 691
    SH3_Eve1_4; Fourth Src homology 3 domain of ADAM-binding protein Eve-1
    cd11818
    Location:711 → 760
    SH3_Eve1_5; Fifth Src homology 3 domain of ADAM-binding protein Eve-1
    PHA03247
    Location:29 → 369
    PHA03247; large tegument protein UL36; Provisional
    cl17036
    Location:415 → 445
    SH3; Src Homology 3 domain superfamily

  3. NM_001128924.2 → NP_001122396.1  SH3 domain-containing protein 19 isoform 3

    See identical proteins and their annotated locations for NP_001122396.1

    Status: REVIEWED

    Description
    Transcript Variant: This variant (3) lacks an in-frame coding exon in the 5' region compared to variant 1. This results in a shorter isoform (3) missing an internal protein segment compared to isoform 1.
    Source sequence(s)
    AW263019, BC032404, BC108893, BM670337, BM831166, CD675494
    Consensus CDS
    CCDS47143.1
    UniProtKB/Swiss-Prot
    Q5HYK7
    Related
    ENSP00000415694.1, ENST00000427414.2
    Conserved Domains (5) summary
    cd11815
    Location:440 → 491
    SH3_Eve1_2; Second Src homology 3 domain of ADAM-binding protein Eve-1
    cd11816
    Location:516 → 566
    SH3_Eve1_3; Third Src homology 3 domain of ADAM-binding protein Eve-1
    cd11817
    Location:606 → 655
    SH3_Eve1_4; Fourth Src homology 3 domain of ADAM-binding protein Eve-1
    cd11818
    Location:675 → 724
    SH3_Eve1_5; Fifth Src homology 3 domain of ADAM-binding protein Eve-1
    cl17036
    Location:379 → 409
    SH3; Src Homology 3 domain superfamily

  4. NM_001243349.2 → NP_001230278.1  SH3 domain-containing protein 19 isoform 2

    See identical proteins and their annotated locations for NP_001230278.1

    Status: REVIEWED

    Description
    Transcript Variant: This variant (4) contains an alternate 5' terminal non-coding exon, and lacks an in-frame coding exon compared to variant 1. This results in a shorter isoform (2, also known as Eve-1b) missing an internal protein segment compared to isoform 1. Variants 2 and 4 encode the same isoform.
    Source sequence(s)
    AK294476, BC045742, BF109395, CF541231
    Consensus CDS
    CCDS47144.1
    UniProtKB/Swiss-Prot
    Q5HYK7
    Related
    ENSP00000423449.1, ENST00000514152.5
    Conserved Domains (6) summary
    cd11815
    Location:476 → 527
    SH3_Eve1_2; Second Src homology 3 domain of ADAM-binding protein Eve-1
    cd11816
    Location:552 → 602
    SH3_Eve1_3; Third Src homology 3 domain of ADAM-binding protein Eve-1
    cd11817
    Location:642 → 691
    SH3_Eve1_4; Fourth Src homology 3 domain of ADAM-binding protein Eve-1
    cd11818
    Location:711 → 760
    SH3_Eve1_5; Fifth Src homology 3 domain of ADAM-binding protein Eve-1
    PHA03247
    Location:29 → 369
    PHA03247; large tegument protein UL36; Provisional
    cl17036
    Location:415 → 445
    SH3; Src Homology 3 domain superfamily

  5. NM_001378121.1 → NP_001365050.1  SH3 domain-containing protein 19 isoform 4

    Status: REVIEWED

    Source sequence(s)
    AC095055, AC104819
    Conserved Domains (6) summary
    cd11814
    Location:699 → 748
    SH3_Eve1_1; First Src homology 3 domain of ADAM-binding protein Eve-1
    cd11815
    Location:779 → 830
    SH3_Eve1_2; Second Src homology 3 domain of ADAM-binding protein Eve-1
    cd11816
    Location:855 → 905
    SH3_Eve1_3; Third Src homology 3 domain of ADAM-binding protein Eve-1
    cd11817
    Location:945 → 994
    SH3_Eve1_4; Fourth Src homology 3 domain of ADAM-binding protein Eve-1
    cd11818
    Location:1014 → 1063
    SH3_Eve1_5; Fifth Src homology 3 domain of ADAM-binding protein Eve-1
    PHA03247
    Location:168 → 649
    PHA03247; large tegument protein UL36; Provisional

  6. NM_001378122.1 → NP_001365051.1  SH3 domain-containing protein 19 isoform 5

    Status: REVIEWED

    Source sequence(s)
    AC095055, AC104819
    Consensus CDS
    CCDS93652.1
    UniProtKB/TrEMBL
    A0A0U1RQE4
    Related
    ENSP00000488951.1, ENST00000604030.7

  7. NM_001378123.1 → NP_001365052.1  SH3 domain-containing protein 19 isoform 6

    Status: REVIEWED

    Source sequence(s)
    AC095055
    Consensus CDS
    CCDS93651.1
    UniProtKB/TrEMBL
    A0A494C1M0
    Related
    ENSP00000499127.1, ENST00000652233.1
    Conserved Domains (6) summary
    cd11814
    Location:640 → 689
    SH3_Eve1_1; First Src homology 3 domain of ADAM-binding protein Eve-1
    cd11815
    Location:720 → 771
    SH3_Eve1_2; Second Src homology 3 domain of ADAM-binding protein Eve-1
    cd11816
    Location:796 → 846
    SH3_Eve1_3; Third Src homology 3 domain of ADAM-binding protein Eve-1
    cd11817
    Location:886 → 935
    SH3_Eve1_4; Fourth Src homology 3 domain of ADAM-binding protein Eve-1
    cd11818
    Location:955 → 1004
    SH3_Eve1_5; Fifth Src homology 3 domain of ADAM-binding protein Eve-1
    PHA03247
    Location:109 → 590
    PHA03247; large tegument protein UL36; Provisional

  8. NM_001378124.1 → NP_001365053.1  SH3 domain-containing protein 19 isoform 7

    Status: REVIEWED

    Source sequence(s)
    AC095055
    Conserved Domains (6) summary
    cd11815
    Location:697 → 748
    SH3_Eve1_2; Second Src homology 3 domain of ADAM-binding protein Eve-1
    cd11816
    Location:773 → 823
    SH3_Eve1_3; Third Src homology 3 domain of ADAM-binding protein Eve-1
    cd11817
    Location:863 → 912
    SH3_Eve1_4; Fourth Src homology 3 domain of ADAM-binding protein Eve-1
    cd11818
    Location:932 → 981
    SH3_Eve1_5; Fifth Src homology 3 domain of ADAM-binding protein Eve-1
    PHA03247
    Location:109 → 590
    PHA03247; large tegument protein UL36; Provisional
    cl17036
    Location:636 → 666
    SH3; Src Homology 3 domain superfamily

  9. NM_001378126.1 → NP_001365055.1  SH3 domain-containing protein 19 isoform 1

    Status: REVIEWED

    Source sequence(s)
    AC095055
    Consensus CDS
    CCDS34077.2
    UniProtKB/Swiss-Prot
    B7Z296, Q08EK1, Q32N10, Q5HYK7, Q5U3B8, Q86XB3, Q8N5E7, Q9UFC8
    Related
    ENSP00000386848.2, ENST00000409252.6
    Conserved Domains (6) summary
    cd11814
    Location:419 → 468
    SH3_Eve1_1; First Src homology 3 domain of ADAM-binding protein Eve-1
    cd11815
    Location:499 → 550
    SH3_Eve1_2; Second Src homology 3 domain of ADAM-binding protein Eve-1
    cd11816
    Location:575 → 625
    SH3_Eve1_3; Third Src homology 3 domain of ADAM-binding protein Eve-1
    cd11817
    Location:665 → 714
    SH3_Eve1_4; Fourth Src homology 3 domain of ADAM-binding protein Eve-1
    cd11818
    Location:734 → 783
    SH3_Eve1_5; Fifth Src homology 3 domain of ADAM-binding protein Eve-1
    PHA03247
    Location:29 → 369
    PHA03247; large tegument protein UL36; Provisional

  10. NM_001378127.1 → NP_001365056.1  SH3 domain-containing protein 19 isoform 1

    Status: REVIEWED

    Source sequence(s)
    AC095055
    Consensus CDS
    CCDS34077.2
    UniProtKB/Swiss-Prot
    B7Z296, Q08EK1, Q32N10, Q5HYK7, Q5U3B8, Q86XB3, Q8N5E7, Q9UFC8
    Conserved Domains (6) summary
    cd11814
    Location:419 → 468
    SH3_Eve1_1; First Src homology 3 domain of ADAM-binding protein Eve-1
    cd11815
    Location:499 → 550
    SH3_Eve1_2; Second Src homology 3 domain of ADAM-binding protein Eve-1
    cd11816
    Location:575 → 625
    SH3_Eve1_3; Third Src homology 3 domain of ADAM-binding protein Eve-1
    cd11817
    Location:665 → 714
    SH3_Eve1_4; Fourth Src homology 3 domain of ADAM-binding protein Eve-1
    cd11818
    Location:734 → 783
    SH3_Eve1_5; Fifth Src homology 3 domain of ADAM-binding protein Eve-1
    PHA03247
    Location:29 → 369
    PHA03247; large tegument protein UL36; Provisional

  11. NM_001378128.1 → NP_001365057.1  SH3 domain-containing protein 19 isoform 2

    Status: REVIEWED

    Source sequence(s)
    AC095055, AC104819
    Consensus CDS
    CCDS47144.1
    Conserved Domains (6) summary
    cd11815
    Location:476 → 527
    SH3_Eve1_2; Second Src homology 3 domain of ADAM-binding protein Eve-1
    cd11816
    Location:552 → 602
    SH3_Eve1_3; Third Src homology 3 domain of ADAM-binding protein Eve-1
    cd11817
    Location:642 → 691
    SH3_Eve1_4; Fourth Src homology 3 domain of ADAM-binding protein Eve-1
    cd11818
    Location:711 → 760
    SH3_Eve1_5; Fifth Src homology 3 domain of ADAM-binding protein Eve-1
    PHA03247
    Location:29 → 369
    PHA03247; large tegument protein UL36; Provisional
    cl17036
    Location:415 → 445
    SH3; Src Homology 3 domain superfamily

  12. NM_001378129.1 → NP_001365058.1  SH3 domain-containing protein 19 isoform 2

    Status: REVIEWED

    Source sequence(s)
    AC095055
    Consensus CDS
    CCDS47144.1
    Conserved Domains (6) summary
    cd11815
    Location:476 → 527
    SH3_Eve1_2; Second Src homology 3 domain of ADAM-binding protein Eve-1
    cd11816
    Location:552 → 602
    SH3_Eve1_3; Third Src homology 3 domain of ADAM-binding protein Eve-1
    cd11817
    Location:642 → 691
    SH3_Eve1_4; Fourth Src homology 3 domain of ADAM-binding protein Eve-1
    cd11818
    Location:711 → 760
    SH3_Eve1_5; Fifth Src homology 3 domain of ADAM-binding protein Eve-1
    PHA03247
    Location:29 → 369
    PHA03247; large tegument protein UL36; Provisional
    cl17036
    Location:415 → 445
    SH3; Src Homology 3 domain superfamily

  13. NM_001378130.1 → NP_001365059.1  SH3 domain-containing protein 19 isoform 2

    Status: REVIEWED

    Source sequence(s)
    AC095055
    Consensus CDS
    CCDS47144.1
    Conserved Domains (6) summary
    cd11815
    Location:476 → 527
    SH3_Eve1_2; Second Src homology 3 domain of ADAM-binding protein Eve-1
    cd11816
    Location:552 → 602
    SH3_Eve1_3; Third Src homology 3 domain of ADAM-binding protein Eve-1
    cd11817
    Location:642 → 691
    SH3_Eve1_4; Fourth Src homology 3 domain of ADAM-binding protein Eve-1
    cd11818
    Location:711 → 760
    SH3_Eve1_5; Fifth Src homology 3 domain of ADAM-binding protein Eve-1
    PHA03247
    Location:29 → 369
    PHA03247; large tegument protein UL36; Provisional
    cl17036
    Location:415 → 445
    SH3; Src Homology 3 domain superfamily

  14. NM_001378131.1 → NP_001365060.1  SH3 domain-containing protein 19 isoform 2

    Status: REVIEWED

    Source sequence(s)
    AC095055, AC104819
    Consensus CDS
    CCDS47144.1
    Conserved Domains (6) summary
    cd11815
    Location:476 → 527
    SH3_Eve1_2; Second Src homology 3 domain of ADAM-binding protein Eve-1
    cd11816
    Location:552 → 602
    SH3_Eve1_3; Third Src homology 3 domain of ADAM-binding protein Eve-1
    cd11817
    Location:642 → 691
    SH3_Eve1_4; Fourth Src homology 3 domain of ADAM-binding protein Eve-1
    cd11818
    Location:711 → 760
    SH3_Eve1_5; Fifth Src homology 3 domain of ADAM-binding protein Eve-1
    PHA03247
    Location:29 → 369
    PHA03247; large tegument protein UL36; Provisional
    cl17036
    Location:415 → 445
    SH3; Src Homology 3 domain superfamily

RefSeqs of Annotated Genomes: GCF_000001405.40-RS_2024_08

The following sections contain reference sequences that belong to a specific genome build. Explain

Reference GRCh38.p14 Primary Assembly

Genomic

  1. NC_000004.12 Reference GRCh38.p14 Primary Assembly

    Range
    151120281..151325605 complement
    Download
    GenBank, FASTA, Sequence Viewer (Graphics)

Reference GRCh38.p14 PATCHES

Genomic

  1. NW_025791772.1 Reference GRCh38.p14 PATCHES

    Range
    57012..262336 complement
    Download
    GenBank, FASTA, Sequence Viewer (Graphics)

Alternate T2T-CHM13v2.0

Genomic

  1. NC_060928.1 Alternate T2T-CHM13v2.0

    Range
    154444161..154649516 complement
    Download
    GenBank, FASTA, Sequence Viewer (Graphics)
Nucleotide Protein
Heading Accession and Version
Protein Accession Links
GenPept Link UniProtKB Link
Q5HYK7.2 GenPept UniProtKB/Swiss-Prot:Q5HYK7

Gene LinkOut

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